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VMD/PhD Alumni Profiles


The VMD-PhD Program has enjoyed a long history of training outstanding veterinarian-scientists. Nothing speaks more to the quality of our graduates than their numerous and diverse accomplishments. When you browse through these profiles, you will see for yourself the value of a Penn VMD/PhD degree.

  • Theresa Alenghat

    Assistant Professor, Cincinnati Children’s Hospital

    • VMD:  2003   Theresa Alenghat, VMD-PhD
    • PhD:  2007
    • Graduate Group:  Cell and Molecular Biology
    • Thesis Topic:  Molecular and physiologic significance of the nuclear receptor corepressor-HDAC3 complex

    After graduating from the program, Dr. Alenghat worked as a Postdoc at Perelman School of Medicine at University of Pennsylvania. She is currently an Assistant Professor, UC Department of Pediatrics at the Cincinnati Children’s Hospital.

  • Stephen Artim

    Resident in Lab Animal Medicine, Division of Comparative Medicine at Stephen Artim, VMD-PhDMassachusetts Institute of Technology

    • VMD: 2015
    • PhD: 2014
    • Graduate Group: Biochemistry and Molecular Biophysics
    • Thesis Topic:  Regulation and Kinase Activity of the TRK Family of Receptor Tyrosine Kinases

    After graduation, Steve moved to Boston to join a LAM residency at MIT.

  • Steven Bensinger

    AsSteven Bensinger, VMD-PhDsociate Professor of Pathology and Laboratory Medicine, Molecular & Medical Pharmacology, University of California at Los Angeles

    • VMD: 1998
    • PhD: 2003
    • Graduate Group: Immunology
    • Thesis Topic: Development and function of a CD4+25+ regulatory T cells

    The Bensinger laboratory is largely interested in understanding how lipid metabolism influences lymphocyte proliferation and adaptive immunity.  

     
  • Jessica Bertout

    Senior Scientist, Presage Biosciences, Inc.Jessica Bertout, VMD-PhD

    • VMD: 2010
    • PhD: 2009
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Hypoxia Inducible Factors Regulate the p53 and Noth Pathways to Promote Cancer

    After graduation, Jessica accepted a postdoctoral position at the Fred Hutchinson Cancer Center in Dr. Jason Biela's laboratory. She is currently at Presage Biosciences, Inc as a Senior Scientist.

  • Susan Bender
    Susan Bender, VMD-PhD

    Pathobiology Lecturer, School of Veterinary Medicine at New Bolton Center of the University of Pennsylvania

    • VMD: 2011
    • PhD: 2010
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Receptor utilization and antiviral CD8 T cell responses during central nervous system infection with a murine coronavirus

    Post-graduation, entered a residency in Anatomic Pathology at the University of Pennsylvania School of Veterinary Medicine. She is currently a Anatomic Pathology Lecturer at University of Pennsylvania.

  • Avrama Blackwell

    Professor of Molecular Neurosciences, George Mason University

    • VMD: 1986
    • PhD: 1988
    • Graduate Group: Bioengineering
    • Thesis Topic: The role of spatial and chromatic parameters in image processing by the visual system

    The Blackwell laboratory studies the molecular, cellular and network mechanisms of long term memory storage using computational and electrophysiological techniques.  The lab studies neuromodulators and their role in Parkinson's disorder and mental illness.

  • Catherine Brinkley
    Dr. Catherine Brinkley, VMD-PhD

    Assistant Professor, Department of Community and Regional Development, University of California at Davis

    • VMD: 2015
    • PhD: 2013
    • Graduate Group: City and Regional Planning
    • Thesis Topic: Fringe Benefits: quantitative and qualitative study of urban form on farmland functions

    Katie is an Assistant Professor at the University of California, Davis.

  • Irene Bukh
    Irene Bukh, VMD-PhD

    Post doctoral position, Perelman School of Medicine, University of Pennsylvania

    • VMD: 2015
    • PhD: 2014
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Blood and Guts: Consequences of Adenovirus Vector Vaccination on T Cell Activation and SIV Susceptibility in Rhesus Macaques

    Post-graduation, Irene is at the Perelman School of Medicine at the Department of Microbiology as a Postdoctoral Researcher.

     

     

  • Sarah Bushmeyer

    Dr. Sarah BushmeyerVeterinarian, Dogwood Pet Hospital

    • VMD: 1992
    • PhD: 1997
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Transcription factor YY1 and its role in regulating transcription of the immunoglobulin light chain enhancer

     

     

     

  • Bryan Cherry
    Bryan Cherry, VMD-PhD

    Interim Director, Regional Epidemiology Program at the Bureau of Communicable Disease Control of the New York State Department of Health

    • VMD: 1997
    • PhD: 2003
    • Graduate Group: Parasitology
    • Thesis Topic: Infection dynamics of Ichthyophthirius multifillis in zebrafish and catfish

    Dr. Cherry's main interest is control and prevention of zoonotic diseases.  Currently, he directs statewide surveillance for communicable diseases occurring in the community setting, evaluates surveillance methods, and implements and evaluates education and prevention programs for communicable diseases.

  • Michelle Cook Sangar
    Michelle Cook Sangar, VMD-PhD

    Director of Pharmacology, Peptide Drug Discovery Program at the Fred Hutchinson Cancer Center

    • VMD: 2011
    • PhD: 2010
    • Graduate Group: Pharmacology
    • Thesis Topic: The role of mitochondrial-targeted human Cytochrome P450 2D6 in drug metabolism and toxicity

    Post-graduation, Michelle accepted a Postdoc position at the Fred Hutchinson Cancer Center in Seattle.  She is now the Director of Pharmacology in the Peptide Drug Discovery Program at Fred Hutchinson. 

  • LaTasha Crawford

    Anatomical Pathology Residency and Post-doctoral Research Position, Johns Hopkins University

    LaTasha Crawford, VMD-PhD
    • VMD: 2011 
    • PhD: 2010 
    • Graduate Group: Neuroscience 
    • Thesis Topic: Probing the Neurophysiology of anxiety: Social stress alters the modulation of serotonin neurons 

    LaTasha's thesis project characterized 2 subpopulations of serotonin cells and revealed that those 2 subpopulations are regulated differently in social stress-based model of anxiety.

     

     

  • Mary Anne Della Fera

    Assistant Research Scientist, University of Georgia

    Chief Scientific Officer, AptoTec, Inc.

    President, Biomedical Information ServicesMary Anne Della Fera, VMD-PhD

    • VMD: 1979
    • PhD: 1980
    • Graduate Group: Anatomy
    • Thesis Topic: Role of CNS cholecystokinin in hypothalamic control of food

    Dr. Della Fera's primary area of interest is in regulation of energy balance of food intake.  In recent years, she has focused on regulation of adipose tissue mass, and, in particular, the effects of natural compounds on adipogenesis, lipolysis, lipogenesis, and adipocyte apoptosis.

  • Lise Desquenne-Clark

    Staff Scientist, The Wistar InstituteLise Desquenne-Clark, VMD-PhD

    • VMD: 1984
    • PhD: 1988
    • Graduate Group: Immunology
    • Thesis Topic: Studies on neonatal transplantation tolerance:  A role for MHC restriction

     

     

     

  • Aimee Edinger

    Associate Professor of Cell and Developmental Biology, University of California at IrvineAimee Edinger, VMD-PhD

    • VMD: 1996
    • PhD: 1999
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: HIV/SIV Entry

    Dr. Edinger's laboratory addresses cancer treatment by trying to exploit metabolic differences to selectively starve cancer cells by down-regulating nutrient transporter proteins.

     

     

  • Duncan Ferguson

    Professor and Department Head or the Department of Comparative Biosciences at the College of Veterinary Medicine, University of Illinois at Urbana-ChampaignDuncan Ferguson, VMD-PhD

    • VMD: 1979
    • PhD: 1982
    • Graduate Group: Pharmacology
    • Thesis Topic: Tyroid hormone metabolism in the isolated perfused rat kidney

    Dr. Ferguson is interested in the interaction of the hypothalamic-pituitary-thyroid (HPT) axis nutrient metabolism and vice versa.

     

     

  • Alan Fine

    Director of the Neuroscience Institute
    Professor in the Department of Physiology and Biophysics, Dalhousie University Faculty of MedicineAlan Fine, VMD-PhD

    • VMD: 1979
    • PhD: 1987
    • Graduate Group: Physiology
    • Thesis Topic: Cortical acetylcholine

    Dr. Fine's laboratory investigates mechanisms of information processing and storage in the central nervous system; development and application of advanced optical imaging methods for neuroscience, including extrinsic and genetically-encoded fluorescent ion and voltage indicators in conjunction with multiphoton, confocal- and fast CCD-imaging.

  • Marnie FitzMaurice Cryer

    Instructor at the College of Veterinary Medicine, Cornell University

    Marnie FitzMaurice Cryer, VMD-PhD

    • VMD: 1996
    • PhD: 2002
    • Graduate Group: Neuroscience
    • Thesis Topic: Role of the superior colliculus in visual detection

     

    Dr. FitzMaurice is the course leader for the Neuroanatomy course at the College of Veterinary Medicine at Cornell University.  She instructs first year classes in topics including anatomy, histology, embryology, physiology and pharmacology.

  • Bruce Freedman

    Associate Professor of Pathobiology and Director of PennVet Imaging Core Facility, School of Veterinary Medicine at the University of PennsylvaniaBruce Freedman, VMD-PhD

    • VMD: 1987
    • PhD: 1992
    • Graduate Group: Physiology
    • Thesis Topic: Expression and function of Kv channels during T cell differentiation

    The general focus of the Freedman laboratory is physiological processes that regulate lymphocyte and macrophage development and function. They are specifically interested in pathways that regulate calcium signaling in T and B cells and in the viral and cellular determinants of viral tropism and pathogenesis.

  • Hannah Galantino-Homer

    Senior Research Investigator for the Laminitis Research Initiative, Department of Clinical Studies at New Bolton Center, School of Veterinary Medicine at the University of PennsylvaniaHannah Galantino-Homer, VMD-PhD

    • VMD: 1993
    • PhD: 2000
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Using molecular genetic techniques to investigate a putative muscarinic receptor homologue in mouse sperm that may be involved in sperm-egg binding and/or the acrosome reaction
  • Brittany Gregory

    Brittany Gregory, VMD-PhDSmall Animal Internship, Wheat Ridge Animal Hospital

    • VMD: 2015
    • PhD: 2013
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Natural variation in the histone demethylase KDM4C, influences cell proliferation through the regulation of specific genes

    Brittany is in Colorado for a SA rotating internship at Wheat Ridge Animal Hospital.  She hopes to then pursue a residency in Pathology.

     

     

  • Joan C Hendricks

    Gilbert S. Kahn Dean of Veterinary Medicine, School of Veterinary Medicine at the University of PennsylvaniaJoan C Hendricks, VMD-PhD

    • VMD: 1980
    • PhD: 1980
    • Graduate Group: Anatomy
    • Thesis Topic: Studies of the effects of small pontine lesions producing paradoxical sleep without atonia in freely behaving cats

    After decades studying physiology and anatomy of sleep and an animal model of sleep apnea (the English bulldog), Dr. Hendricks turned her attention to a simpler model - Drosophila.  She showed, using behavioral, pharmaceutical, and molecular approaches, that Drosophila's rest state has sleep-like properties.  Dr. Hendricks' lab used this model organism to show that the cAMP-PKA-CREB signaling pathway is involved in rest regulation as well as its well-known role in long-term memory consolidation.

  • Alice Hsu

    Veterinarian, Blackhorse Pike Animal HospitalAlice Hsu, VMD-PhD

    • VMD: 2009
    • PhD: 2008
    • Graduate Group: Immunology
    • Thesis Topic: Leishmania mexicana infection

    The goal of Alice's project was to better understand how Leishmania mexicana infection induces a non-healing response in mice that are fully capable of controlling infection with L. major via the generation of a strong cell-mediated Th1 response.

  • Richard Jimenez
    Richard Jimenez, VMD-PhD

    Small Animal Rotating Internship, VCA Animal Hospital 

    • VMD: 2015
    • PhD: 2016
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Mechanisms of translational control in early embryonic development

    Post-graduation, Richard will be doing a Small Animal Rotating Internship at a VCA hospital in Maryland near Washington DC.

     

  • Douglas Jones

    Professor, College of Veterinary Medicine at Iowa State UniversityDouglas Jones, VMD-PhD

    • VMD: 1989
    • PhD: 1993
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Defensin expression in the human small intestine:  A study of the human defensin family

    Dr. Jones studies Leishmania amazonesis, a disease that can have a variety of symptoms from skin ulcers and organ disease to facial disfigurations and immune system malfunctions.

     

     

     

  • Colleen Kane

    Principal Scientist, Janssen BiotechColleen Kane, VMD-PhD

    • VMD: 2007
    • PhD: 2007
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Helminth antigens modulate TLR-ligand induced dendritic cell activation

  • James Kehler

    Dr. James KehlerDirector of Scientific Programs and Alliances, New York Stem Cell Foundation

    • VMD: 2002
    • PhD: 2004
    • Graduate Group: Cell and Molecular Biology - Genetics and Gene Regulation
    • Thesis Topic: Investigating the role of Oct4, a transcription factor, during germline development of the mouse

     

     

     

  • Robert Kieval

    Executive Vice President and Chief Technology Officer, CVRx, Inc.Robert Kieval, VMD-PhD

    • VMD: 1987
    • PhD: 1991
    • Graduate Group: Physiology
    • Thesis Topic: Electrical cell coupling in post-ischemic, arrhythmia-prone myocardium

    Dr. Kieval is the founder and Chief Technology Officer of CVRx, a private company located in Minneapolis, Minnesota. The company has developed a proprietary, electrical stimulation-based implantable medical devise for the treatment of hypertension that cannot be controlled with medications. This product is market-approved in Europe and is in advanced clinical trials in the United States.

  • Catrina King
    Catrina King, VMD-PhD

    Senior Manager of Research Animal Facility and Veterinary Services, Regeneron

    • VMD: 2012
    • PhD: 2011
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Micro-RNA in colorectal malignancies

    Catrina's thesis entitled "LIN28B, let-7, and the molecular pathogenesis of colon cancer" explored the role of proteins that regulate microRNAs in colon tumors, and suggests that these factors may serve as prognostic markers for humans diagnosed with colon cancer. After graduation, Catrina was a Resident in laboratory animal medicine at Columbia University. Catrina is currently at Regeneron as a Senior Manager of Research.

  • Joyce Knoll

    Associate Professor and Chair of the Department of Biomedical Science, Cummings School of Veterinary Medicine at Tufts UniversityJoyce Knoll, VMD-PhD

    • VMD: 1984
    • PhD: 1987
    • Graduate Group: Pathology
    • Thesis Topic: A mouse model for alpha thalassemia:  A characterization using electron microscopy, autoradiography, and iron kinetics

    Dr. Knoll is interested in veterinary hematology, cytology, and clinical endocrinology.


  • Debra Kotloff

    Veterinarian, Oak Tree Veterinary HospitalDebra Kotloff, VMD-PhD

    • VMD: 1984
    • PhD: 1987
    • Graduate Group: Immunology
    • Thesis Topic: The regulation of isotype expression on T dependent antibody responses

     

     

     

  • David Kowalczyk

    Director of Regulatory Affairs, Monsanto CompanyDavid Kowalczyk, VMD-PhD

    • VMD: 1975
    • PhD: 1976
    • Graduate Group: Pharmacology
    • Thesis Topic: The effect of Pb on K flux in frog sartorius muscle
  • Dara Kraitchman

    Professor of Radiology and Radiological Science, School of Medicine at Johns Hopkins UniversityDara Kraitchman, VMD-PhD

    • VMD: 1992
    • PhD: 1996
    • Graduate Group: Bioengineering
    • Thesis Topic: Integrated magnetic resonance imaging studies of cardiac function and perfusion

    Dr. Kraitchman's research intersts are concentrated on non-invasive imaging of cardiovascular disease.  She has been actively involved in developing new methods to image myocardial function and perfusion using magnetic resonance imaging (MRI).

  • Erika Lin-Hendel
    Erika Lin-Hendel, VMD-PhD

    Staff Scientist, Biomin

    • VMD: 2015
    • PhD: 2016
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Control of mitochondrial trafficking in neurons

    Post-graduation, Erika will be working in agricultural research and global food animal production and food security for the Austrian company, Biomin.

     

     

     

  • Leslie MacGregor-Levine

    Formerly Chief Intellectual Property Counsel, Perkin-ElmerLeslie MacGregor-Levine, VMD-PhD

    • VMD: 1981
    • PhD: 1986
    • Graduate Group: Physiology
    • Thesis Topic: Biochemical and electrophysical abnormalities in retinas of diabetic animals
  • Katherine Masek Hammerman

    Principal Scientist and Pathologist, Pfizer Inc.Katherine Masek Hammerman, VMD-PhD

    • VMD: 2002
    • PhD: 2006
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Innate immunity to Toxoplasma gondii 

    After graduation Katie entered a Postdoctoral Fellow/Resident position in Comparative and Experimental Pathology at the New England Primate Research Center, Harvard University. She is currently at Pfizer as a Principal Scientist and Pathologist.

  • Amy Matthews
    Amy Matthews, VMD-PhD

    Veterinarian, Frontier Wellness Center

    • VMD: 2001
    • PhD: 2001
    • Graduate Group: Immunology
    • Thesis Topic: The role of B lymphocytes in murine hepatitis (strain A59 clearance and pathology)

     

     

     

     

  • James MacLeod

    Professor of Veterinary Science, John S. and Elizabeth A. Knight Chair and Director of the Equine Initiative, University of Kentucky

    James MacLeod, VMD-PhD
    • VMD:  1984
    • PhD:  1990
    • Graduate Group:  Pathology
    • Thesis Topic:  Hormonal regulation of hepatic P450 monooxygenases

    Dr. MacLeod's laboratory studies biological and biomedical aspects of the musculoskeletal system, with an emphasis on the growth and maturation of articular cartilage, the development of osteoarthritis, repair of cartilage lesions, and the effects of intra-articular glucocorticoid medications on chrondrocyte function.

  • Patricia McManus

    Formerly, Clinical Pathologist at IDEXX Reference Laboratories

    Associate Editor of Veterinary Clinical Pathology, An International Journal of Laboratory MedicinePatricia McManus, VMD-PhD

    • VMD: 1980
    • PhD: 1983
    • Graduate Group: Pathology
    • Thesis Topic: Bone marrow failure syndromes

    Dr. McManus handles daily pathology cases and occasional resident training at IDEXX. Dr. McManus has been editing for the Veterinary Clinical Pathology journal for four years. She first started out as a Section Editor for Hematology and Immunology, and then recently became an Associate Editor. She is also a member of the IDEXX Medical Review Board.

  • Albee Messing

    Professor of Neuropathology in the Department of Comparative Biosciences, School of Veterinary Medicine at the University of Wisconsin at MadisonAlbee Messing, VMD-PhD

    • VMD: 1978
    • PhD: 1982
    • Graduate Group: Pathology
    • Thesis Topic: Alpha-bungarotoxin receptors of cultured chick ciliary ganglion neurons

    Research in Dr. Messing's laboratory is directed at understanding developmental and pathologic aspects of glial cell biology in the nervous system of the mouse, with a particular focus on astrocytes and their major intermediate filament protein, GFAP.

  • Natalie Miller

    Natalie Miller, VMD-PhDVeterinary Medical Officer, US Food and Drug Administration

    • VMD: 2013
    • PhD: 2012
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Population genetics of T. gondii

    Post-graduation, Natalie worked as a veterinarian in private practice. Natalie is currently at the FDA as a Veterinary Medical Officer reviewing pre-clinical trial data.

     

     

  • Richard Miselis

    Formerly Professor of Animal Biology

    Head of the Laboratories of Anatomy, School of Veterinary Medicine at the University of PennsylvaniaRichard Miselis, VMD-PhD

    • VMD: 1973
    • PhD: 1973
    • Graduate Groups: Biology
    • Thesis Topic: The glucoprivic control of ingestive behavior

    The Miselis laboratory was interested in the neurobiology of visceral function and ingestive behavior.

  • Jaime Modiano

    Perlman Professor of Oncology and Comparative Medicine

    Director of Animal Cancer Center and Research Program, College of Veterinary Medicine at the University of MinnesotaJaime Modiano, VMD-PhD

    • VMD: 1991
    • PhD: 1991
    • Graduate Group: Immunology
    • Thesis Topic: Calcium and protein kinase C as mediators of T cell activation

    Jaime Modiano is the Director of the University of Minnesota Animal Cancer Care and Research Program. His lab is interested in understanding the basic differences between normal cells and tumor cells.

     

     

  • Kenneth Mohn

    Senior Associate Director of Technology Assessment, Boehringer Ingelheim Vetmedica Inc.Kenneth Mohn, VMD-PhD

    • VMD: 1987
    • PhD: 1991
    • Graduate Group: Molecular Biology/Genetics
    • Thesis Topic: Characterization of the immeidate-early growth response in regenerating liver and insulin-stimulated Reuber H-35 cells

    Dr. Mohn's career to date has focused on discovery and/or acquisition of novel pharmaceuticals for virtually the entire range of economically important medical conditions in domestic species.

  • Stephanie Murphy

    Director of the Division of Comparative Medicine, Office of Research Infrastructure Programs in the Division of Program Coordination, and Strategic Initiatives of the Office of the Director at the National Institutes of HealthStephanie Murphy, VMD-PhD

    • VMD: 1994
    • PhD: 1996
    • Graduate Group: Biochemistry
    • Thesis Topic: Possible mechanisms of neurotoxicity and dopamine-mediated injury in the newborn brain during hypoxia and posthypoxic reoxygenation

    Dr. Murphy is responsible for programs at NIH that provide funding for the development and access to critical animal models, which offer essential clues to a broad range of human disorders such as Parkinson’s disease, diabetes, multiple sclerosis, and AIDS. She provides scientific leadership for strengthening and enhancing a critical part of the biomedical research continuum, providing support for animal models that build the bridge between basic science and human medicine, with discoveries in one species enhancing understanding of another. She provides funding for pre- and post-doctoral training and career development for veterinary scientists who provide unique expertise to translational biomedical research, and establishes collaborations between DCM-supported grantees and grantees funded by the NIH Institutes and Centers. Her office sets scientific priorities through the development of long-term strategic plans and execution of funding decisions, and manages the direction of operations and activities through the implementation of NIH policies and resource allocation.

  • Xuan Pan

    Assistant Professor in the Department of Medical Sciences, University of WisconsinXuan Pan, VMD-PhD

    • VMD: 2008
    • PhD: 2010
    • Thesis Topic: YY1 functions in B lymphocyte development

    After graduating from the program, Xuan and her husband moved to Michigan where she pursued a small animal clinical internship at Michigan State University.  After completion of this program she took an oncology residency position at University of Wisconsin. She is currently an Assistant Professor of Oncology at the University of Wisconsin working on the function of transcription factor YY1. 

     

     

  • Thomas Parsons

    Thomas Parsons, VMD-PhDAssociate Professor of Clinical Studies at New Bolton Center, School of Veterinary Medicine at the University of Pennsylvania

    Associate Professor of Otorhinolaryngology, School of Medicine at the University of Pennsylvania

    • VMD: 1986
    • PhD: 1989
    • Graduate Group: Neuroscience
    • Thesis Topic: Optical monitoring of the electrical activity of neuronal ensembles constructed from identified invertebrate cells

     

     

  • Nikkita Patel

    Independent ContractorNikkita Patel, VMD-PhD

    • VMD: 2012
    • PhD: 2015
    • Graduate Group: Epidemiology and Biostatistics 
    • Thesis Topic: Characterization of Illegal Wildlife Trade Networks

    Nikki is currently an independent contractor with the Wildlife Conservation Society.

     

     

     

  • Caroline Patten

    Owner/Veterinarian, Liberty Veterinary ClinicCaroline Patten, VMD-PhD

    • VMD: 2009
    • PhD: 2007
    • Graduate Group: Neuroscience
    • Thesis Topic: Intracellular signaling pathways of the melanocortin type 3 and type 4 receptors

     

     

     

     

  • Jenni Punt

    Professor  & Associate Dean for Student Research in the Department of Pathology and Cell Biology, College of Physicians and Surgeons at Columbia UniversityJenni Punt, VMD-PhD

    • VMD: 1988
    • PhD: 1991
    • Graduate Group: Immunology
    • Thesis Topic: Regulation of positive selection and lineage commitment during thymocyte development

    Dr. Punt is an immunologist who has worked at multiple levels of education and research (high school, college, professional school). 

     

     

  • Mark Pykett

    President and CEO, Agilis Biothearpeutics LLCMark Pykett, VMD-PhD

    • VMD:  1991
    • PhD:  1994
    • Graduate Group:  Cell and Molecular Biology
    • Thesis Topic:  Identification of molecular alterations in human meningiomas

     

     

     

     

  • Sarah Ralston

    Professor in the Department of Animal Sciences at the School of Environmental and Biological Sciences, Rutgers, the State University of New JerseySarah Ralston, VMD-PhD

    • VMD: 1980
    • PhD: 1982
    • Graduate Group: Anatomy
    • Thesis Topic: Factors in the control of feed intake in ponies

    Dr. Ralston is board certified by the American College of Veterinary Nutrition and has served on the Board of directors for many years. Her primary research focus throughout her career has been on the effect of rations on glucose and insulin metabolism in horses. 


  • Gregory Rak

    Research Investigator, Bristol-Myers SquibbGregory Rak, VMD-PhD

    • VMD: 2012
    • PhD: 2011
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Granule secretion through the actin network at the lytic NK cell synapse

    After graduation, Gregory worked as a Postdoctoral Fellow at the Perelman School of Medicine at University of Pennsylvania. He currently works at Bristol-Myers Squibb as a Research Investigator.


  • Laurel Redding

    Laurel Redding, VMD-PhDPostdoctoral Position, School of Veterinary Medicine at the University of Pennsylvania

    • VMD: 2012
    • PhD: 2015
    • Graduate Group: Epidemiology and Biostatistics
    • Thesis Topic: Understanding the Use of Antibiotics on Small Dairy Farms in the Developing World

    Laurel is currently doing postdoctoral work with Drs. Gary Smith and Shelly Rankin at the University of Pennsylvania, School of Veterinary Medicine.

  • Peter Bart Reiner

    Professor, Kinsmen Laboratory of Neurological Research at the University of British ColumbiaPeter Bart Reiner, VMD-PhD

    • VMD: 1982
    • PhD: 1984
    • Graduate Group: Anatomy
    • Thesis Topic: The activity of neurons in the area of the Locus Coeruleus during the states of sleep

    Dr. Reiner has studied the neurobiology of behavioral states and the molecular underpinnings of neurodegenerative disease. He also has experience in the private sector as President and CEO of Active Pass Pharmaceuticals, a drug discovery company that he founded to tackle the Alzheimer's disease.


  • John Robertson

    Professor of Biomedical Engineering in the Department of Basic Science, Virginia Tech School of Medicine and Research InstituteJohn Robertson, VMD-PhD

    • VMD: 1976
    • PhD: 1977
    • Graduate Group: Pathology
    • Thesis Topic: Tubulointestinal and glomerulonephritis in the Brown Norway rat:  A model for autoimmune nephritis

    Dr. Robertson's research interests include comparative oncology of malignant melanoma and malignant lymphoma, chronic renal disease, and implantable biomaterials.

  • Mary Robinson

    Director of the Penn Vet Equine Pharmacology Laboratory in the Department of Clinical Studies at New Bolton Center, School of Veterinary Medicine at the University of PennsylvaniaMary Robinson, VMD-PhD

    • VMD: 2010
    • PhD: 2009
    • Thesis Topic: Changes in Oxygen Tension Rapidly and Reversibly Regulate Macrophage Nitric Oxide Production

     

     

     

  • Allen Rushmer

    Veterinarian and Owner, Next Generation Embryo TransferAllen Rushmer, VMD-PhD

    • VMD: 1975
    • PhD: 1981
    • Graduate Group: Physiology
    • Thesis Topic: Examination of the interaction of oocytes and their surrounding cumulus cells

  • Melissa Sanchez

    Pathologist, Antech DiagnosticsMelissa Sanchez, VMD-PhD

    • VMD: 2002
    • PhD: 2006
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Characterization of neutralizing antibodies against West Nile virus in mice and horses

     

     

     

     

  • Eric Sandgren

    Associate Professor of Experimental Pathology, School of Veterinary Medicine at the University of Wisconsin at MadisonEric Sandgren, VMD-PhD

    • VMD: 1986
    • PhD: 1992
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Transforming growth factor alpha and multistage cancer in the mouse

    The overall goal of Dr. Sandgren's research program is to understand genetic and extracellular mechanisms that regulate cell and tissue growth, whether that growth is a reparative response to tissue damage or an abnormal condition leading to cancer.

  • Linda Schuler

    Professor of Comparative Biosciences, School of Veterinary Medicine at the University of Wisconsin at MadisonLinda Schuler, VMD-PhD

    • VMD: 1981
    • PhD: 1980
    • Graduate Group: Physiology
    • Thesis Topic: Cholesterol metabolism in the ovary of the rat

    The Schuler lab is interested prolactin and related hormones and the determinants of cell and context specific actions, including ligand specificity, receptor signal transduction, receptor processing and cross talk with other signaling pathways.

  • W. Douglas Sheffield

    Executive Director of Clinical Affairs, Spiration, IncW. Douglas Sheffield, VMD-PhD

    • VMD: 1975
    • PhD: 1977
    • Graduate Group: Pathology
    • Thesis Topic: Pathogenesis of experimental autoimmune encephalomyelitis

    Dr. Sheffield has over 25 years experience directing the research and development of medical devices, biologies and pharmaceutical products.  Previously, he was Director of New Technologies at Northstar Neuroscience, Inc., as well as at Johnson & Johnson where he was Ethicon's Director of Pharmaceutical Research and Development.  He also served as Director of Surgical R&D at Ethicon Endo-Surgery, leading to the development of numerous minimally invasive procedures.  Dr. Sheffield is an inventor with multiple patents relating to medical devices and has authored and co-authored numerous medical publications.


  • Spencer Shelly

    Spencer Shelly, VMD-PhDVeterinarian, Private Practice 

    • VMD: 2012
    • PhD: 2013
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: "Functional analysis of the Herpes Simplex Virus glycoprotein B membrane proximal domain." 

    After graduation, Spencer gained clinical experience in general or emergency practice. In the future, he is interested in pursuing a career in public health aspects of infectious disease.

     

     

  • Bruce Smith

    Professor of Pathology and Scientist at the Scott-Ritchey Research Center, College of Veterinary Medicine at Auburn UniversityBruce Smith, VMD-PhD

    • VMD: 1988
    • PhD: 1993
    • Graduate Group: Genetics
    • Thesis Topic: Canine phosphofructokinase deficiency

    Dr. Smith's research program in muscular dystrophies focuses on identifying canine models of these diseases and applying novel genetic therapies to their treatment.

     

     

  • Gail Smith

    Professor of Orthopedic Surgery, School of Veterinary Medicine at the University of PennsylvaniaGail Smith, VMD-PhD

    • VMD: 1974
    • PhD: 1982
    • Graduate Group: Materials Science Engineering
    • Thesis Topic: Passivation and corrosion kinetics of Stainless Steel 316L orthopaedic implants

    With a background in physical sciences, Dr. Smith has explored and innovated materials and mechanics applicable to orthopedics. Having a reliable quantitative measure of hip laxity allows accurate prediction of dogs and cats that are susceptible to hip OA later in life.  This technology is known as "PennHIP" and is now being adopted worldwide.


  • John Stambaugh

    Associate Veterinarian & Chief of Surgery, Boston Road Animal HospitalJohn Stambaugh, VMD-PhD

    • VMD: 1974
    • PhD: 1975
    • Graduate Group: Physiology
    • Thesis Topic: The effects of calcification on diffusion in the epiphysical plate

     

     

     

     

  • Todd Strochlic

    Todd Strochlic, VMD-PhDAssistant Professor in the Department of Biochemistry & Molecular Biology, Drexel University 

    • VMD: 2008
    • PhD: 2008
    • Graduate Group: Cell and Molecular Biology - Cell Biology and Physiology
    • Thesis Topic: Regulated protein sorting in the endosomal system

    After graduating, Dr. Strochlic was a Postdoctoral Fellow at the Fox Chase Cancer Center.  He is currently an Assistant Professor at Drexel University.

  • Ruth Sullivan

    Research Scientist and Lecturer in the Laboratory for Optical and Computational Instrumentation at the McPherson Eye Research Institute, University of Wisconsin at MadisonRuth Sullivan, VMD-PhD

    • VMD: 1991
    • PhD: 1996
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Conotruncal cardiac malformation and neural tube defects in transgenic mice expressing a dominant negative CS43/ß-galactosidase fusion protein

     

     

     

  • Steven Suter

    Steven Suter, VMD-PhDAssociate Professor of Medical Oncology and Medical Director for the Canine Bone Marrow Transplant Unit, College of Veterinary Medicine at North Carolina State University

    • VMD: 1995
    • PhD: 2000
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: The isolation and characterization of a population of canine putative hematopoietic progenitor cells and their use as targets for retroviral mediated gene therapy

    The Suter lab studies the molecular underpinnings of canine and feline lymphoma and is involved in the development of novel therapeutics for these diseases.

  • James Thomson

    Director of Regenerative Biology, Morgridge Institute for Research

    Professor, University of WisconsinJames Thomson, VMD-PhD

    • VMD: 1985
    • PhD: 1988
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Understanding genetic imprinting in early mammalian development

    Dr. Thomson directed the group that reported the first isolation of embryonic stem cell lines from a non-human primate in 1995, work that led his group to the first successful isolation of human embryonic stem cell lines in 1998.  In November 2007 Dr. Thomson's team succeeded in isolating similar pluripotent stem cells from human somatic cells.  Dr. Thomson was elected into the National Academy of Sciences in 2008 and is the recipient of the 2011 King Faisal International Prize, and the recipient of the 2011 Albany Medical Center Prize in Medicine and Biomedical Research.


  • Alexander Travis

    Associate Professor of Reproductive Biology and Wildlife Conservation and Director of the Cornell Center for Wildlife Conservation, Baker Institute for Animal Health at Cornell UniversityAlexander Travis, VMD-PhD

    • VMD: 1995
    • PhD: 1999
    • Graduate Group: Cell and Molecular Biology
    • Thesis Topic: Characterization, targeting and functional analyses of male germ cell-specific isoforms of type I hexokinase in the mouse

    Dr. Travis' laboratory primarily studies the compartmentalization of signaling and metabolic pathways in male germ cells. 

    Dr. Travis was a recipient of the NIH Pioneer Award in 2009.


  • Susan Volk

    Assistant Professor of Small Animal Surgery, School of Veterinary Medicine at the University of Pennsylvania

    • Susan Volk, VMD-PhD, Penn VetVMD: 1995
    • PhD: 1998
    • Graduate Group: Pathology
    • Thesis Topic: Bone morphogenetic protein signaling during chondrocyte maturation

    Dr. Volk’s research interests focus on defining regulatory cues important to direct progenitor cell homing to sites of injury, as well as, in vitro and in vivo signals important to direct their differentiation into appropriate target cell types for use in tissue engineering strategies in human and veterinary medicine.

  • John Wolfe

    Professor of Pathology, School of Veterinary Medicine at the University of Pennsylvania

    Director, Walter Flato Goodman Center for Comparative Medical Genetics

    Associate Director, Core Center for Gene Therapy of Cystic Fibrosis and Other Genetic Disease

    Stokes Investigator, Children's Hospital of PhiladelphiaJohn Wolfe, VMD-PhD

    • VMD: 1982
    • PhD: 1986
    • Graduate Group: Immunology 
    • Thesis Topic: H-2-linked control of retrovirus production in Gross and Friend virus-induced tumor cell lines.

    Dr. Wolfe's laboratory studies methods for gene and stem cell therapies in the central nervous system. They use animal homologues of human genetic diseases as test systems. The approaches include: ex vivo gene transfer using retrovirus and lentivirus vector-modified neural stem cells transplanted to the brain; and direct injection of herpesvirus, adeno-associated virus, and lentivirus vectors.